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Zanthoxylum dipetalum var. dipetalum

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Zanthoxylum

Description

Key Characters:

Growth Form: Trees 4–15 m tall.

Stems: Trunk up to 30 cm in diameter, bark finely fissured–reticulate, new growth glabrous or finely puberulent to tomentose, young branches subtended by a pair of bud scales.

Roots:

Leaves: Leaves compound. Alternate. Leaflets 3–7, thick, coriaceous, lowest pair strongly reduced, (0.2–)1.5–3(–4) cm long, rarely 1 or both reduced leaflets absent, normal leaflets 6–15(–36) cm long, (2.5–)3.5–9(–13.5) cm wide, lateral petiolules articulated only at base, 3–9 mm long, terminal petiolule twice articulated, at base and near leaflet, 8–30 mm long; lowest pair of leaflets stipelliform or, if expanded, then usually folded or strongly revolute, arising immediately below lowest pair of normal leaflets, normal leaflets oblong–elliptic, elliptic to broadly elliptic or sometimes broadly obovate. Leaflet apices obtuse to rounded. Base cuneate to rounded. Surfaces glabrous; conspicuously glandular punctate. Margins entire rarely weakly emarginate. Normal leaflets lateral veins usually 6–8 pairs, prominent, higher order venation forming a reticulate pattern. Rachis terete (cylindrical) or winged. Stipules absent.

Flowers: Flowers usually 5–15 in terminal or leaf–opposed, open, cymose inflorescences usually 5–10 cm long, peduncles 10–40 mm long, pedicels 3–6(–8) mm long, subtended by a pair of bud scales; bracteoles 1 mm or less long. Flowers unisexual, actinomorphic. Calyx of 4 sepals, broadly ovate–deltate, 1–1.5 mm long, ciliolate. Corolla of 2 or sometimes 4 petals; petals white or yellowish white, sometimes tinged reddish, lanceolate to ovate, 6–10 mm long. Stamens 4, alternate with the petals, glabrous or pubescent, absent in pistillate flowers; anthers medifixed, dithecal. Ovary superior, 2-carpellate, but usually only 1 carpel developing, placentation axile, rudimentary in staminate flowers; ovules 2 per carpel, pendulous; style short, ± coherent; stigmas obliquely orbicular, capitate.

Fruit: Follicles 1–2; ovoid; 15–25(–33) mm long; glandular–pitted; apex conspicuously beaked; the beak 3–8 mm long; exocarp rugose; endocarp coriaceous. Seeds usually 1 per follicle; on a slender funicle; black; ovoid; 10–26 mm long.

Ploidy:

Habitat: Dry to mesic forest; sometimes along margins of wet forest.

Elevation Range: 260–1,280 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Other References

Wagner et al. 1990:1214 (K, O, Mo, H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Zanthoxylum dipetalum var. dipetalum hvnp, kipuka puaulu. next to nps transect roadside PRESERVED_SPECIMEN drake, d. collector number: k.m. 19 Hawaii BISH 3/25/1988
2 Zanthoxylum dipetalum var. dipetalum waioli-waipa ridge, 10 ft. e. of crest Dry hillside PRESERVED_SPECIMEN christensen, c. collector number: 195 Oahu BISH 4/27/1977
3 Zanthoxylum dipetalum var. dipetalum kohua ridge Forest PRESERVED_SPECIMEN degener, o. degener, i.; hansen, h.w. collector number: 23920 Kauai BISH 9/9/1955
4 Zanthoxylum dipetalum var. dipetalum kaholuamanu, mountains PRESERVED_SPECIMEN forbes, c.n. collector number: 347 k Kauai BISH 9/1/1909
5 Zanthoxylum dipetalum var. dipetalum pupukea-kahuku road, kahuku PRESERVED_SPECIMEN st.john, h. collector number: 25972 Oahu BISH 10/28/1956
6 Zanthoxylum dipetalum var. dipetalum kaholuamano PRESERVED_SPECIMEN rock, j.f.c. collector number: 10209 Oahu BISH 10/1/1911
7 Zanthoxylum dipetalum var. dipetalum pupukea - kahuku trail, paunalu - waimea ridge. koolau range PRESERVED_SPECIMEN st.john, h. collector number: 11585 Oahu BISH 3/6/1932
8 Zanthoxylum dipetalum var. dipetalum kaholuamano Open hillside. PRESERVED_SPECIMEN rock, j.f.c. collector number: 10209 Oahu BISH 10/1/1911
9 Zanthoxylum dipetalum var. dipetalum pupukea trail PRESERVED_SPECIMEN swezey, o.h. collector number: s.n. Oahu BISH 6/16/1935
10 Zanthoxylum dipetalum var. dipetalum oio, paumalu trail forest PRESERVED_SPECIMEN degener, o. collector number: 17724 Oahu BISH 6/16/1940
11 Zanthoxylum dipetalum var. dipetalum waimea canyon rim, below road, 11.8 mi fm waimea PRESERVED_SPECIMEN marr, k.l. collector number: km 9 Kauai BISH 3/26/1988
12 Zanthoxylum dipetalum var. dipetalum puaulu PRESERVED_SPECIMEN rock, j.f.c. collector number: 10208 Hawaii BISH 7/1/1911
13 Zanthoxylum dipetalum var. dipetalum kokee, awaawapuhi trail PRESERVED_SPECIMEN stone, b.c. collector number: 3337 Kauai BISH 4/12/1960
14 Zanthoxylum dipetalum var. dipetalum oio, paumalu trail PRESERVED_SPECIMEN degener, o. collector number: 17590 French Frigate Shoals BISH 6/16/1940
15 Zanthoxylum dipetalum var. dipetalum pohakuao, hanging valley between kalalau and hanakoa Lowland mesic forest PRESERVED_SPECIMEN wood, k.r. perlman, s.; lau, j. collector number: 1769 Kauai BISH 4/2/1992
16 Zanthoxylum dipetalum var. dipetalum waianae mts., puu kaua to pohakea pass, 3rd ridge from summit, ne facing PRESERVED_SPECIMEN marr, k.l. palmer, d.d. collector number: 54 Oahu BISH 6/24/1988
17 Zanthoxylum dipetalum var. dipetalum 3 mi. down makaha ridge road, n facing slope PRESERVED_SPECIMEN flynn, t. randall, j. collector number: 1810 Kauai BISH 7/15/1986
18 Zanthoxylum dipetalum var. dipetalum kipuka puaulu PRESERVED_SPECIMEN rock, j.f.c. collector number: 13019 Hawaii BISH 8/8/1917
19 Zanthoxylum dipetalum var. dipetalum oahu PRESERVED_SPECIMEN st.john, h. collector number: 11585 Oahu BISH 3/6/1932
20 Zanthoxylum dipetalum var. dipetalum waimea, kauai PRESERVED_SPECIMEN mann, h. brigham, w.t. collector number: 581 Kauai BISH
21 Zanthoxylum dipetalum var. dipetalum kumuweia ridge, e of waineke swamp Forest PRESERVED_SPECIMEN degener, o. collector number: 21551 Kauai BISH 1/8/1952
22 Zanthoxylum dipetalum var. dipetalum kokee. app. 30m past kokee state park sign on right side of main road. Kokee State Park PRESERVED_SPECIMEN marr, k.l. collector number: k.m. 15 Kauai BISH 3/26/1988
23 Zanthoxylum dipetalum var. dipetalum ridge above ekahanui gulch Schinus terebinthifolius PRESERVED_SPECIMEN takeuchi, w.n. pyle, l. collector number: 2063 Oahu BISH 3/10/1985
24 Zanthoxylum dipetalum var. dipetalum olokele (?) valley PRESERVED_SPECIMEN lydgate, j.m. Kauai BISH 11/1/1911
25 Zanthoxylum dipetalum var. dipetalum awaawapuhi trail; honopu PRESERVED_SPECIMEN lane, i.e. collector number: 56-625 Kauai BISH 12/29/1956
26 Zanthoxylum dipetalum var. dipetalum puu kaua, e ridge Exocarpos; Myrsine, Dubautia, Pelea, Metrosideros PRESERVED_SPECIMEN marr, k.l. palmer, d.d. collector number: 52 Oahu BISH 6/24/1988
27 Zanthoxylum dipetalum var. dipetalum bird park (kipuka puaulu) PRESERVED_SPECIMEN morley, h. collector number: 104-h Oahu BISH 8/11/1934
28 Zanthoxylum dipetalum var. dipetalum kaholuamano PRESERVED_SPECIMEN rock, j.f.c. collector number: 5201 Kauai BISH 9/28/1909
29 Zanthoxylum dipetalum var. dipetalum kaunala gulch, kaunala, koolau range, kahuku forest res., lower woods lower woods PRESERVED_SPECIMEN st.john, h. hosaka, e.y. collector number: 20164 Oahu BISH 2/9/1941
30 Zanthoxylum dipetalum var. dipetalum puu kaua, waianae mts., honouliuli mucky stagnant pond in trail to ohia dieback plot 24; rainforest; with Juncus, Eleocharis, and Hypericum PRESERVED_SPECIMEN st.john, h. collector number: 12226 Hawaii BISH 11/6/1932
31 Zanthoxylum dipetalum var. dipetalum kaholuamano, near mountain house PRESERVED_SPECIMEN rock, j.f.c. collector number: 5540 Kauai BISH 9/28/1909
32 Zanthoxylum dipetalum var. dipetalum waianae range, pu'u ka'ala, above waialua on the duponte trail deep woods PRESERVED_SPECIMEN b. c. stone 3185 Kauai BISH 12/20/1959
33 Zanthoxylum dipetalum var. dipetalum kaholuamano Occasional in weedy disturbed places on pyroclastic soil PRESERVED_SPECIMEN rock, j.f.c. collector number: 5201 Hawaii BISH 9/28/1909
34 Zanthoxylum dipetalum var. dipetalum ~30 m past kokee sign, main road PRESERVED_SPECIMEN marr, k.l. collector number: 15 Kauai BISH 3/26/1988
35 Zanthoxylum dipetalum var. dipetalum haw. natl. park PRESERVED_SPECIMEN fagerlund, g.o. mitchell, a.l. collector number: 827 Hawaii BISH
36 Zanthoxylum dipetalum var. dipetalum waianae mts., honouliuli fr, ridge to puu kaua exposed arid cliff PRESERVED_SPECIMEN herbst, d.r. obata, j. collector number: 5263 Oahu BISH 3/16/1975
37 Zanthoxylum dipetalum var. dipetalum nuuanu valley PRESERVED_SPECIMEN hillebrand, w.b. collector number: s.n. Oahu BISH
38 Zanthoxylum dipetalum var. dipetalum kahuku mauka shrubby forest PRESERVED_SPECIMEN degener, o. et al. collector number: 19544 Oahu BISH 3/20/1949
39 Zanthoxylum dipetalum var. dipetalum lava tube region, kilauea PRESERVED_SPECIMEN morley, h. collector number: 104-h Hawaii BISH 8/6/1934
40 Zanthoxylum dipetalum var. dipetalum waimea canyon rim, 11.8 mi fm waimea, 50 m below road On ridge. PRESERVED_SPECIMEN marr, k.l. collector number: km 44 Oahu BISH 6/13/1988
41 Zanthoxylum dipetalum var. dipetalum pupukea paumala fr, koolau mts., jeep road PRESERVED_SPECIMEN spence, g. collector number: 8 BISH 10/31/1971
42 Zanthoxylum dipetalum var. dipetalum waimea district; kawaiiki zone, below kalukaula ridge, on ridge looking down into drainage of koaie canyon and north of waialae. Degraded Metrosideros - Acacia koa Diverse Mesic Forest Myoporum, Cryptocarya, Kadua affinis, Alyxia, Claoxylon sandwicensis, Dodonea, Leptocaphylla, Dryopteris, Zanthoxylum, Dubautia, Psychotria mariniana, Scaevola procera, Melicope PRESERVED_SPECIMEN tangalin, n. kishida, w. collector number: 2573 Kauai BISH 3/23/2011
43 Zanthoxylum dipetalum var. dipetalum koolau range, pupukea-kahuku trail, in small ravine near road PRESERVED_SPECIMEN pearsall, g.a. collector number: 522 Oahu BISH 5/18/1960
44 Zanthoxylum dipetalum var. dipetalum kalalau, very back on e side of valley, 50 ft up talus slope on e side of n running gulch, gulch just w of the mainstream. Old relic mixed mesic forest with weedy understory. Christella parasitica, Nephrolepis, Bryophyllum pinnatum, Adenophorus hispidulum, Elephantopus mollis, midstory mostly open with Lantana and Psydrax, loose open canopy of Aleurites moluccana, Melia azedarach PRESERVED_SPECIMEN tangalin, n. kishida, w. collector number: 2385 Kauai BISH 8/22/2010
45 Zanthoxylum dipetalum var. dipetalum kumuwela ridge, waimea Dry open woods PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13835 Kauai BISH 12/28/1933
46 Zanthoxylum dipetalum var. dipetalum honopu, awaawapuhi valley Mesic forest, canopy ht ave 55 ft PRESERVED_SPECIMEN stone, b.c. collector number: 1592 Kauai BISH 12/29/1956
47 Zanthoxylum dipetalum var. dipetalum kalalau, on a ridge at the back of the valley Cultivated area running north from garden entrance along edge of forest. PRESERVED_SPECIMEN hobdy, r.w. collector number: 1815 Maui BISH 8/20/1983
48 Zanthoxylum dipetalum var. dipetalum hind ranch, between puuwaawaa and puu iki Rain forest PRESERVED_SPECIMEN hatheway, w.h. collector number: 452 Molokai BISH 11/20/1950
49 Zanthoxylum dipetalum var. dipetalum waianae mts., ekahnanui gulch, ridge, e puu kaua Myrsine emarginata, Myrsine sandwicensis, Exocarpos gaudichaudii, Styphelia, Dubautia, Plantaginea, Pittosporum flocculosum, Scaevola gaudichaudiana PRESERVED_SPECIMEN perlman, s.p. obata, j. collector number: 5100 Oahu BISH 1/15/1987
50 Zanthoxylum dipetalum var. dipetalum kokee, awaawapuhi trail Forest PRESERVED_SPECIMEN stone, b.c. collector number: 3338 Kauai BISH 4/12/1960
51 Zanthoxylum dipetalum var. dipetalum Makaleha Mtns., Kapaahi, End of Kahuna Rd, past water tank, along stream and up along East Ridge, then East along back bowl and down ridge just to the east of trail ridge. Lowland Wet Metrosideros-Dicranopteris windward Forest. PRESERVED_SPECIMEN Natalia Tangalin 268 Kauai PTBG 1/28/2005
52 Zanthoxylum dipetalum var. dipetalum waimea district, kawaiula valley, kuia natural area reserve. PRESERVED_SPECIMEN k. r. wood & m. query 14967 A 5/9/2012
53 Zanthoxylum dipetalum var. dipetalum PRESERVED_SPECIMEN O Degener|F Salucop|V Arlantico AK
54 Zanthoxylum dipetalum var. dipetalum waimea district, koaie, headwater gulches and stream, well above koaie river, north of mohihi-waialae trail Shady bank PRESERVED_SPECIMEN k. r. wood, m. kirkpatrick & s. p. perlman 15970 MIN 6/24/2014
55 Zanthoxylum dipetalum var. dipetalum kaunala gulch, kaunala, koolau range, kahuku forest reserve PRESERVED_SPECIMEN h. st. john 20164 MNHN 2/9/1941
56 Zanthoxylum dipetalum var. dipetalum PRESERVED_SPECIMEN h. mann jr. 581 NY
57 Zanthoxylum dipetalum var. dipetalum hanalei district, upper limahuli valley, downstream from base camp u drainage on west side. PRESERVED_SPECIMEN n. tangalin 898 NY 8/8/2006
58 Zanthoxylum dipetalum var. dipetalum kipukapuaulu, near kilauea on slopes of mauna loa. PRESERVED_SPECIMEN j. f. rock 10208 Hawaii US
59 Zanthoxylum dipetalum var. dipetalum Koaie, headwater gulches and stream, well above Koaie river, north of Mohihi-Waialae trail. Metrosideros transitional mesic to wet forest and riparian vegetation PRESERVED_SPECIMEN K. R. Wood 15970 Kauai PTBG 6/24/2014
60 Zanthoxylum dipetalum var. dipetalum honopu trail Beside trail PRESERVED_SPECIMEN marr, k.l. nakamura, j. collector number: km 41 Kauai BISH 6/11/1988
61 Zanthoxylum dipetalum var. dipetalum waianae range, puu kaala, above waialua on the duponte ridge trail at 2800 ft. alt Diverse, mixed wet-mesic native forest PRESERVED_SPECIMEN stone, b.c. collector number: 3185 Kauai BISH 12/20/1959
62 Zanthoxylum dipetalum var. dipetalum w waimea drainage basin PRESERVED_SPECIMEN forbes, c.n. collector number: 970 k Kauai BISH 7/3/1917
63 Zanthoxylum dipetalum var. dipetalum waimea canyon, mile 11.3, hwy. 550 Zanthoxylum kauaense, Pelea anisata, Myrsine, Acacia; dry PRESERVED_SPECIMEN flynn, t. lorence, d.; de lappe, r. collector number: 2212 Kauai BISH 5/21/1987
64 Zanthoxylum dipetalum var. dipetalum kipuka puaulu Moist Aleurites forest. PRESERVED_SPECIMEN russ, g.w. Kauai BISH 6/1/1930
65 Zanthoxylum dipetalum var. dipetalum kaholuamano In full sun. PRESERVED_SPECIMEN rock, j.f.c. collector number: 2106 Oahu BISH 3/3/1909
66 Zanthoxylum dipetalum var. dipetalum hvnp, kipuka puaulu. next to nps transect One other large individual and at least 5 other saplings in Kipuka Puaulu. Three saplings and one seedling beneath this tree. PRESERVED_SPECIMEN drake, d. collector number: k.m. 19 Hawaii BISH 3/25/1988
67 Zanthoxylum dipetalum var. dipetalum ridge, puu kaua cliff edge PRESERVED_SPECIMEN takeuchi, w.n. et al. collector number: waianae 126 Oahu BISH 7/15/1984
68 Zanthoxylum dipetalum var. dipetalum waimea canyon, 2 mi. e of kokee ranger station, road to molilii, 1st stream crossing PRESERVED_SPECIMEN henrickson, j.s. collector number: 4056 Kauai BISH 8/1/1969
69 Zanthoxylum dipetalum var. dipetalum na pali survey, supp. sta. c., back of valley on e side Diverse mesic forest on steep slope PRESERVED_SPECIMEN perlman, s.p. hobdy, b.; gon, s. collector number: 10374 Kauai BISH 6/13/1989
70 Zanthoxylum dipetalum var. dipetalum kipuka vuaulu PRESERVED_SPECIMEN j. f. rock 10208 A 7/20/1911
71 Zanthoxylum dipetalum var. dipetalum kipuka vuaulu In neglected garden PRESERVED_SPECIMEN j. f. rock 10208 IND 7/20/1911
72 Zanthoxylum dipetalum var. dipetalum Lahaina District , Maui, Kauaula Valley, south slope. About one-third down from valley rim towards stream, northwest aspect PRESERVED_SPECIMEN H. Oppenheimer H21405 NY 2/13/2014
73 Zanthoxylum dipetalum var. dipetalum PRESERVED_SPECIMEN H. Mann Jr. 581 NY
74 Zanthoxylum dipetalum var. dipetalum puaulu Cultivated PRESERVED_SPECIMEN j. f. c. rock 10208 NY 7/7/1911
75 Zanthoxylum dipetalum var. dipetalum lahaina district , maui, kauaula valley, south slope. about one-third down from valley rim towards stream, northwest aspect PRESERVED_SPECIMEN h. oppenheimer h21405 NY 2/13/2014
76 Zanthoxylum dipetalum var. dipetalum KAUAI: Kokee, off Milolii ridge road in Acacia koa- Metrosideros mesic forest with Xylosma hawaiiense, Alphitonia ponderosa, Pleomele aurea, Psychotria mariniana, Psidium cattleianum PRESERVED_SPECIMEN Steve Perlman 19479 Kauai PTBG 3/29/2005
77 Zanthoxylum dipetalum var. dipetalum Waianae Mts., Honouliuli, North Palawai gulch, north facing slope. In Acacia koa-Metrosideros lowland mesic forest. PRESERVED_SPECIMEN Steve Perlman 6159 Oahu PTBG 7/12/1987
78 Zanthoxylum dipetalum var. dipetalum KAUAI: Limahuli valley,off ridge between Limahuli and Hanakapiai valleys, 1950 ft. elev. on ridge crest in Metrosideros-Dicranopteris linearis wet forest w/ Antidesma, Ilex, Bobea, Syzygium sandwicensis, Psychotria, Smilax, Hedyotis terminalis, Freycinetia, Pleomele, Diospyros, Myrsine,Pipturus, Nestegis sandwicensis, Bidens, Alyxia Boehmeria grandis, Cibotium, Rubus rosifolius, Clidemia PRESERVED_SPECIMEN Steve Perlman 16671 Kauai PTBG 6/22/1999
79 Zanthoxylum dipetalum var. dipetalum Kawaiula Valley, Kuia Natural Area Reserve Metrosideros-Acacia mesic forest PRESERVED_SPECIMEN K. R. Wood 14967 Kauai PTBG 5/9/2012
80 Zanthoxylum dipetalum var. dipetalum Upper Limahuli Preserve, near upper weatherport. Wainiha side. Wet drainage. PRESERVED_SPECIMEN Merlin Edmonds ME76 Kauai PTBG 1/25/2012
81 Zanthoxylum dipetalum var. dipetalum HAWAII: Puuwaawaa near SW corner and ESE of Puu Iki; degraded lowland dry forest with Myrsine lanaiensis, Sophora chrysophylla, Chamaesyce olowaluana. Threats: cattle, ants, Pennisetm setaceaum, Grevelia robusta. Elev. 3470 PRESERVED_SPECIMEN K. R. Wood 7767 Hawaii PTBG 4/7/1999
82 Zanthoxylum dipetalum var. dipetalum Koaie Canyon, 1 mile east of Lonomea camp. Closed to open Metrosideros-Diospyros mixed mesic forest with riparian vegetation. PRESERVED_SPECIMEN K. R. Wood 12457 Kauai PTBG 7/4/2007
83 Zanthoxylum dipetalum var. dipetalum Waiakoali headwaters, south branch. Metrosideros-Dicranopteris-Diplopterygium pinnatum, Cheirodendron montane mixed wet forest and riparian vegetation PRESERVED_SPECIMEN K. R. Wood 17450 Kauai PTBG 6/27/2017
84 Zanthoxylum dipetalum var. dipetalum Upper Limahuli Valley, down stream from base camp up drainage on W side. Metrosideros-Dicranopteris wet forest. PRESERVED_SPECIMEN Natalia Tangalin 898 Kauai PTBG 8/9/2006
85 Zanthoxylum dipetalum var. dipetalum Nuololo, North facing slopes above northern branch. Metrosideros-Acacia montane mesic forest. PRESERVED_SPECIMEN K. R. Wood 15098 Kauai PTBG 7/18/2012
86 Zanthoxylum dipetalum var. dipetalum KAUAI: Pohakuao, eastern drainages below Manono Ridge, relictual Diospyros-Netrosideros mesic forest with Rauvolfia sandwicensis, Pleomele aurea, Streblus pendulinus, Psychotri mariniana, Tetraplasandra waimeae, Microlepia strigosa, Canavalia napaliensis, Carex wahuensis, C. meyenii; Threats: goats, Lantana camara, Pluchea carolinensis, Erigeron karvinskianus, Kalanchoe pinnata. Elev. 1450 ft. PRESERVED_SPECIMEN K. R. Wood 8055-A Kauai PTBG 11/17/1999
87 Zanthoxylum dipetalum var. dipetalum Kawaiiki valley. Lowland Diospyros mixed mesic summer dry forest PRESERVED_SPECIMEN K. R. Wood 16102 Kauai PTBG 9/23/2014
88 Zanthoxylum dipetalum var. dipetalum Koholuamanu behind Waimea. PRESERVED_SPECIMEN C. N. Forbes 347a.K Kauai PTBG 9/1/1909
89 Zanthoxylum dipetalum var. dipetalum Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. PRESERVED_SPECIMEN K. R. Wood 13822 Kauai PTBG 7/16/2009
90 Zanthoxylum dipetalum var. dipetalum KAUAI: Kalalau, between first and second Kalalau lookout, Metrosideros montane mesic forest with Zanthoxylum dipetalum, Pouteria sandwicensis, Pritchardia minor, Xylosma hawaiiense, Exocarpos luteolus, Psychotria greenwelliae, Understory: Diplazium sandwichianum, Ground Cover: Carex meyenii PRESERVED_SPECIMEN K. R. Wood 11128 Kauai PTBG 2/8/2005
91 Zanthoxylum dipetalum var. dipetalum Mahanaloa Valley, Above Weller fence #2 and West. Diverse Mesic Forest PRESERVED_SPECIMEN Natalia Tangalin 1956 Kauai PTBG 3/6/2009
92 Zanthoxylum dipetalum var. dipetalum Poopooiki Valley, Kuia Natural Area Reserve. Metrosideros-Acacia mixed mesic forest PRESERVED_SPECIMEN K. R. Wood 16294 Kauai PTBG 2/5/2015
93 Zanthoxylum dipetalum var. dipetalum KAUAI: Mahanaloa valley, north aspect in diverse mesic forest with Diospyros hillebrandii, Hedyotis terminalis, Pisonia, Diplazium sandwichianum, Doodia kunthiana, Aleurites moluccana, Psidium cattleianum, Rubus rosifolius, Lantana camara, Acacia koa, Metrosideros, Carex wahuensis PRESERVED_SPECIMEN K. R. Wood 5017 Kauai PTBG 3/1/1996
94 Zanthoxylum dipetalum var. dipetalum KAUAI: Honopu; upper north fork headwaters; follow dividing ridge of Kalalau & Honopu; 3800 ft contour south to gulch; follow gulch WSW; Metrosideros-Acacia-Dicranopteris Mesic Forest w/ Dodonaea, Cheirodendron, Melicope anisata, Zantho- xylum dipetalum, Psychotria hexandra, P. mariniana, P. gree- nwellii, Diplazium, Microlepia, Doodia. Threats: Pigs, Deer, Rubus rosifolius, Erigeron, Lonicera. Elev. 3500-3700 ft. PRESERVED_SPECIMEN K. R. Wood 4975 Kauai PTBG 2/21/1996
95 Zanthoxylum dipetalum var. dipetalum Plateau near end of road to Waiakoali headwaters, central branch. Metrosideros-Cheirodendron montane wet, steep south facing closed to open forest. PRESERVED_SPECIMEN K. R. Wood 14458 Kauai PTBG 1/21/2011
96 Zanthoxylum dipetalum var. dipetalum Kalalau, very back of lower valley, forested slopes below and east of Pihea. Diospyros mixed mesic forest. PRESERVED_SPECIMEN K. R. Wood 10899 Kauai PTBG 8/11/2004
97 Zanthoxylum dipetalum var. dipetalum Kalalau valley, drainage west of Pohakulele Camp and east of Huna. Diospyros mixed mesic forest. PRESERVED_SPECIMEN K. R. Wood 11655 Kauai PTBG 12/15/2005
98 Zanthoxylum dipetalum var. dipetalum Kawaiiki Ridge, above Koaie twin falls. Metrosideros-Acacia mesic forest. PRESERVED_SPECIMEN K. R. Wood 13857 Kauai PTBG 8/1/2009
99 Zanthoxylum dipetalum var. dipetalum KAUAI: Honopu, north facing relic forest along steep precipitous slopes; Metrosideros-Acacia mesic forest with Psychotria mariniana, Melicope anisata, Melicope barbigera, Pouteria sandwicensis, Dodonaea viscosa, Dianella sandwicensis, Pritchardia minor, Tetraplasandra waimeae. PRESERVED_SPECIMEN K. R. Wood 7729 Kauai PTBG 3/22/1999
100 Zanthoxylum dipetalum var. dipetalum W. Maui, Kauaula Valley, south slope. About one-third down from valley rim towards stream, northwest aspect. PRESERVED_SPECIMEN Hank Oppenheimer H21405 Maui PTBG 2/13/2014
101 Zanthoxylum dipetalum var. dipetalum HAWAII: Puuwaawaa near SW corner and ESE of Puu Iki; degraded lowland dry forest with Myrsine lanaiensis, Sophora chrysophylla, Chamaesyce olowaluana. Threats: cattle, ants, Pennisetm setaceaum, Grevellia robusta. Elev. 3500 aa lava with large surface boulders and rich brown soil PRESERVED_SPECIMEN K. R. Wood 7769 Hawaii PTBG 4/7/1999
102 Zanthoxylum dipetalum var. dipetalum Kawaiiki Ridge, slope above Koaie, West of twin falls. Metrosideros-Acacia montane mixed mesic forest, summer dry. PRESERVED_SPECIMEN K. R. Wood 15418 Kauai PTBG 2/17/2013
103 Zanthoxylum dipetalum var. dipetalum Kawaiiki Zone, Below Kaluakaula Ridge, on ridge looking down into drainage of Koaie Canyon and north of Waialae. Degraded Metrosideros - Acacia koa Diverse Mesic Forest PRESERVED_SPECIMEN Natalia Tangalin 2573 Kauai PTBG 3/23/2011
104 Zanthoxylum dipetalum var. dipetalum Lihue-Koloa Forest Reserve. Ridge trail from Kahili Mountain Park, SW gulch from main ridge trail, one ridge E of Tetraplasandra bisattenuata # 1. Metrosideros-Dicranopteris linearis lowalnd Wet Forest. PRESERVED_SPECIMEN Natalia Tangalin 1485 Kauai PTBG 1/12/2008
105 Zanthoxylum dipetalum var. dipetalum Honopu Rim. Acacia koa - Metrosideros polymorpha - Dicranopteris linearis montane mesic forest PRESERVED_SPECIMEN K. R. Wood 16533 Kauai PTBG 7/10/2015
106 Zanthoxylum dipetalum var. dipetalum KAUAI: Plateau just S of Awaawapuhi stream head. Diverse mesic forest intermixed with open patches of Dicranopteris. With Metrosideros, Ilex, Scaevola sp., Dodonea, Pouteria, Psychotria greenwelliae, Melicope anisata, Coprosma, Psidium cattleianum, Carex meyenii, Alyxia stellata, and occaisional Hedychium. PRESERVED_SPECIMEN Clay Trauernicht 750 Kauai PTBG 2/19/2009
107 Zanthoxylum dipetalum var. dipetalum KAUAI: *Waimea District, Koai`e Canyon, 1 mile east of Lonomea camp, *closed to open Metrosideros-Diospyros mixed mesic forest with riparian vegetation, associated w/ Cheirodendron fauriei, Antidesma platyphylla, Pouteria sandwicensis, Pleomele aurea, Dodonaea viscosa, Pritchardia minor, understory ferns of Microlepia strigosa, Diplazium sandwichianum, Dryopteris fusco-atra, PRESERVED_SPECIMEN K. R. Wood 12459 Kauai PTBG 7/4/2007
108 Zanthoxylum dipetalum var. dipetalum HAWAII: Puuwaawaa near SE corner; 120 deg. and 100 m SE of cabin; north of road; lowland forest with Myrsine, Sophora chrysophylla, Chamaesyce olowaluana. Threats: cattle, ants, Pennisetm setaceaum, Grevellia robusta. Elev. 3670 aa lava with large surface boulders and rich brown soil PRESERVED_SPECIMEN K. R. Wood 7771 Hawaii PTBG 4/7/1999
109 Zanthoxylum dipetalum var. dipetalum Kuia Valley; north facing forested slopes. Diverse mesic forest. PRESERVED_SPECIMEN K. R. Wood 7803 Kauai PTBG 4/29/1999
110 Zanthoxylum dipetalum var. dipetalum Haupu summit region; *open shrubland/herbland with sparse tree cover of Metrosideros polymorpha; with Dicranopteris linearis, Cibotium glaucum, Sadleria pallida along with groupings of native trees such as Hedyotis terminalis; Psychotria mariniana; Antidesma platyphyllum var. hillebrandii; Bobea brevipes; Xylosma hawaiiense; Cheirodendron fauriei; Melicope feddei; and Tetraplasandra spp. PRESERVED_SPECIMEN K. R. Wood 11973 Kauai PTBG 7/14/2006
111 Zanthoxylum dipetalum var. dipetalum Hikimoe, north facing slopes below road, small patches of Diospyros sandwicensis forest with occasional Metrosideros polymorpha, Coprosma waimeae, Psychotria greenwelliae, Zanthoxylum dipetalum, Psydrax odorata, Pouteria sandwicensis, Nestegis sandwicensis, Pleomele aurea, Pisonia sp., Syzygium sandwicensis, gulch is mostly dominated by secondary invasive species PRESERVED_SPECIMEN K. R. Wood 14226 Kauai PTBG 6/1/2010
112 Zanthoxylum dipetalum var. dipetalum Puu Waawaa ranch, Kaupulehu, SE of Puu Waawaa hill, below Delissea cinder cone, near to cabin, north facing. Dry forest. PRESERVED_SPECIMEN Steve Perlman 17434 Hawaii PTBG 2/28/2001
113 Zanthoxylum dipetalum var. dipetalum Awaawapuhi trail, GPS coordinates approximate PRESERVED_SPECIMEN Anna Sugiyama AS002 Kauai PTBG 2/11/2020
114 Zanthoxylum dipetalum var. dipetalum OAHU: Puu Kaua Region, Lualualei, north facing cliff along ridge just to north of Kaua, dominated with non-native Schinus-Melinus with Carex wahuensis, Carex meyenii, Luzula hawaiiensis, Eragrostis grandis, Lipochaeta tenuis, Dianella sandwicensis, Chamaesyce multiflora, Pittosporum confertiflorum, Zanthoxylum dipetalum, Alyxia oliviformis, Schiedea hookeri, Schiedea ligustrina, Lepidium arbuscula,, Artemisia australis, Peperomia tetraphylla, Panicum beecheyi, Polypodium pellucidum, PRESERVED_SPECIMEN K. R. Wood 10645 Oahu PTBG 3/31/2004
115 Zanthoxylum dipetalum var. dipetalum KAUAI: Koai'e Stream, up side-stream SE at 3000 ft, on north facing banks, Metrosideros mixed mesic transitional wet, Myrsine linearifolia, Bobea timonioides, Ilex anomala, Zanthoxylum dipetalum, Claoxylon sandwicensis, Tetraplasandra spp PRESERVED_SPECIMEN K. R. Wood 7664 Kauai PTBG 2/3/1999
116 Zanthoxylum dipetalum var. dipetalum PRESERVED_SPECIMEN w. l. stern & s. carlquist 1316 Kauai US
117 Zanthoxylum dipetalum var. dipetalum Waimea District, Koaie, headwater gulches and stream, well above Koaie river, north of Mohihi-Waialae trail Transitional mesic to wet forest and riparian vegetation PRESERVED_SPECIMEN Wood, K. R.; Kirkpatrick, M.; Perlman, S. P. 15970 Kauai US 6/24/2014
118 Zanthoxylum dipetalum var. dipetalum Waimea Canyon; 2 miles E of Kokee ranger station and Museum along road to Molilii at first stream crossing where flume begins. PRESERVED_SPECIMEN Henrickson, J. S. 4056 Kauai US 8/1/1969
119 Zanthoxylum dipetalum var. dipetalum lahaina district, w. maui, kauaula valley, south slope, about 1/3 down from valley rim towards stream. PRESERVED_SPECIMEN h. l. oppenheimer, k. m. bustamente & s. p. perlman h21405 Maui US 2/13/2014
120 Zanthoxylum dipetalum var. dipetalum kumuwela ridge PRESERVED_SPECIMEN o. degener & i. degener 28094 Kauai US 9/14/1955
121 Zanthoxylum dipetalum var. dipetalum lahaina district, west maui, kauaula valley, below helu summit, on west facing slope Mesic forest PRESERVED_SPECIMEN s. p. perlman, h. l. oppenheimer & m. pagit 24927 Maui US 4/13/2016
122 Zanthoxylum dipetalum var. dipetalum Vicinity forest ranger station at Kokee, Pipeline trail PRESERVED_SPECIMEN Stern, W. L.; Carlquist, S. 1316 Kauai US 8/1/1961
123 Zanthoxylum dipetalum var. dipetalum vicinity forest ranger station at kokee, pipeline trail PRESERVED_SPECIMEN w. l. stern & s. carlquist 1316 Kauai US
124 Zanthoxylum dipetalum var. dipetalum Kipukapuaulu, near Kilauea on slopes of Mauna Loa. PRESERVED_SPECIMEN Rock, J. F. 10208 Hawaii US 7/11/2020
125 Zanthoxylum dipetalum var. dipetalum Kaholuamanu, behind Waimea PRESERVED_SPECIMEN Forbes, C. N. 347aK. Kauai US 9/9/2020
126 Zanthoxylum dipetalum var. dipetalum Kumuwela Ridge PRESERVED_SPECIMEN Degener, O.; Degener, I. 28094 Kauai US 9/14/1955
127 Zanthoxylum dipetalum var. dipetalum Makai of Puu Moa PRESERVED_SPECIMEN Degener, O.; Salucop, --; Arlantico 11926 Oahu US 4/3/1938
128 Zanthoxylum dipetalum var. dipetalum Kumuweia Ridge E of Waineke Swamp PRESERVED_SPECIMEN Degener, O.; Greenwell, A. B. 21551 Kauai US 1/8/1952
129 Zanthoxylum dipetalum var. dipetalum kaholuamanu, behind waimea PRESERVED_SPECIMEN c. n. forbes 347ak. Kauai US
130 Zanthoxylum dipetalum var. dipetalum puaulu. PRESERVED_SPECIMEN j. f. rock 10208 US
131 Zanthoxylum dipetalum var. dipetalum lahaina district, kauaula valley, south slope, below helu summit, west maui. PRESERVED_SPECIMEN s. p. perlman, h. l. oppenheimer & k. m. bustamente 23854 Kauai US 2/13/2014
132 Zanthoxylum dipetalum var. dipetalum Mountains in vicinity of Kaholuamanu. PRESERVED_SPECIMEN Forbes, C. N. 347.K Kauai US 9/9/2020
133 Zanthoxylum dipetalum var. dipetalum kilauea, kipuka puaulu PRESERVED_SPECIMEN j. f. rock 5916 Hawaii US
134 Zanthoxylum dipetalum var. dipetalum Hanalei District, Upper Limahuli Valley, downstream from base camp u drainage on west side. Wet forest PRESERVED_SPECIMEN Tangalin, N. 898 Kauai US 8/8/2006
135 Zanthoxylum dipetalum var. dipetalum oio, paumalu trail PRESERVED_SPECIMEN o. degener 17205 Oahu US 6/16/1940
136 Zanthoxylum dipetalum var. dipetalum Puaulu. PRESERVED_SPECIMEN Rock, J. F. 10208 US 7/11/2020
137 Zanthoxylum dipetalum var. dipetalum Lahaina District, West Maui, Kauaula Valley, below Helu summit, on west facing slope Mesic forest PRESERVED_SPECIMEN S. P. Perlman, H. L. Oppenheimer & M. Pagit 24927 Maui US 4/13/2016
138 Zanthoxylum dipetalum var. dipetalum waimea canyon; 2 miles e of kokee ranger station and museum along road to molilii at first stream crossing where flume begins. PRESERVED_SPECIMEN j. s. henrickson 4056 Oahu US 8/1/1969
139 Zanthoxylum dipetalum var. dipetalum haupu summit region, steep upper southwestern ridges. PRESERVED_SPECIMEN k. r. wood & m. wood 11973 Hawaii US 7/14/2006
140 Zanthoxylum dipetalum var. dipetalum hind ranch, between puuwaawaa and puu iki PRESERVED_SPECIMEN w. h. hatheway 452 Hawaii US 11/20/1950
141 Zanthoxylum dipetalum var. dipetalum koolau range, pupukea-kahuku trail, paumalu-waimea ridge. PRESERVED_SPECIMEN h. st. john 11585 Hawaii US 3/6/1932
142 Zanthoxylum dipetalum var. dipetalum kahuku mauka PRESERVED_SPECIMEN o. degener, h. j. lam, h. clay, r. saiki & t. murashige 19544 Oahu US 3/20/1949
143 Zanthoxylum dipetalum var. dipetalum Lahaina District, Kauaula Valley, south slope, below Helu summit, West Maui. Mesic forest PRESERVED_SPECIMEN Perlman, S. P.; Oppenheimer, H. L.; Bustamente, K. M. 23854 Maui US 2/13/2014
144 Zanthoxylum dipetalum var. dipetalum na pali survey, supp. sta. c, in back of valley on e side, on steep slope PRESERVED_SPECIMEN s. p. perlman, b. hobdy & s. gon 10374 Kauai US 6/13/1989
145 Zanthoxylum dipetalum var. dipetalum Kokee. Awa'awapuhiu trail. In forest. PRESERVED_SPECIMEN Stone, B. C. 3338 Kauai US 4/12/1960
146 Zanthoxylum dipetalum var. dipetalum Kalalau, between first and second Kalalau lookout. PRESERVED_SPECIMEN Wood, K. R. 11128 Kauai US 2/8/2005
147 Zanthoxylum dipetalum var. dipetalum Awaawapuhi Trail, Honopu PRESERVED_SPECIMEN Lane, I. E. 56-625 Kauai US 12/29/1956
148 Zanthoxylum dipetalum var. dipetalum Waimea District, Nuololo, north facing slopes above northern branch. Montane mesic forest. PRESERVED_SPECIMEN K. R. Wood, M. Query & M. Kirkpatrick 15098 Kauai US 7/18/2012
149 Zanthoxylum dipetalum var. dipetalum Kilauea, Kipuka Puaulu PRESERVED_SPECIMEN Rock, J. F. 5916 Hawaii US 12/14/2020
150 Zanthoxylum dipetalum var. dipetalum puu kaua, waianae mts. PRESERVED_SPECIMEN h. st. john 20125 Kauai US 12/1/1940
151 Zanthoxylum dipetalum var. dipetalum koolau range, pupukea-kahuku trail, insmall ravine near road PRESERVED_SPECIMEN g. pearsall 522 Oahu US 6/18/1960
152 Zanthoxylum dipetalum var. dipetalum Waimea District, Koaie, headwater gulches and stream, well above Koaie river, north of Mohihi-Waialae trail Transitional mesic to wet forest and riparian vegetation PRESERVED_SPECIMEN K. R. Wood, M. Kirkpatrick & S. P. Perlman 15970 Kauai US 6/24/2014
153 Zanthoxylum dipetalum var. dipetalum Hind ranch, between Puuwaawaa and Puu Iki PRESERVED_SPECIMEN Hatheway, W. H. 452 Hawaii US 11/20/1950
154 Zanthoxylum dipetalum var. dipetalum Kaunala Gulch, Kaunala, Koolau Range, Kahuku Forest Reserve PRESERVED_SPECIMEN St. John, H. 20164 Oahu US 2/9/1941
155 Zanthoxylum dipetalum var. dipetalum Kahuku mauka PRESERVED_SPECIMEN Degener, O.; Lam, H. J.; Clay, H. F.; Saiki, R.; Murashige, T. 19544 Oahu US 3/20/1949
156 Zanthoxylum dipetalum var. dipetalum kalalau valley, drainage west of pohakulele camp and east of huna. Mixed mesic forest PRESERVED_SPECIMEN k. r. wood & m. demotta 11655 Kauai US 12/15/2005
157 Zanthoxylum dipetalum var. dipetalum Puu Kaua, Waianae Mts. PRESERVED_SPECIMEN St. John, H. 12226 Oahu US 11/6/1932
158 Zanthoxylum dipetalum var. dipetalum Koolau Range, Pupukea-Paumalu Forest Reserve, Pupukea Road, slope below trail PRESERVED_SPECIMEN Cowan, R. S. 774 Oahu US 1/25/1948
159 Zanthoxylum dipetalum var. dipetalum Na Pali Survey, Supp. Sta. C, in back of valley on E side, on steep slope PRESERVED_SPECIMEN Perlman, S. P.; Hobdy, B.; Gon, S. 10374 Kauai US 6/13/1989
160 Zanthoxylum dipetalum var. dipetalum kaholuamano PRESERVED_SPECIMEN a. s. hitchcock 15296 Kauai US 10/20/1916
161 Zanthoxylum dipetalum var. dipetalum Lahaina District, West Maui, Kauaula Valley, below Helu summit, on west facing slope Mesic forest PRESERVED_SPECIMEN Perlman, S. P.; Oppenheimer, H. L.; Pagit, M. 24927 Maui US 4/13/2016
162 Zanthoxylum dipetalum var. dipetalum kumuweia ridge e of waineke swamp PRESERVED_SPECIMEN o. degener & a. b. greenwell 21551 Kauai US 1/8/1952
163 Zanthoxylum dipetalum var. dipetalum Waimea District, Nuololo, north facing slopes above northern branch. Montane mesic forest. PRESERVED_SPECIMEN Wood, K. R.; Query, M.; Kirkpatrick, M. 15098 Kauai US 7/18/2012
164 Zanthoxylum dipetalum var. dipetalum Lahaina District, W. Maui, Kauaula Valley, south slope, about 1/3 down from valley rim towards stream. PRESERVED_SPECIMEN H. L. Oppenheimer, K. M. Bustamente & S. P. Perlman H21405 Maui US 2/13/2014
165 Zanthoxylum dipetalum var. dipetalum awa`awapuhi trail PRESERVED_SPECIMEN Kauai US
166 Zanthoxylum dipetalum var. dipetalum awaawapuhi trail, honopu PRESERVED_SPECIMEN i. lane 56-625 Kauai US 12/29/1956
167 Zanthoxylum dipetalum var. dipetalum Halemanu. PRESERVED_SPECIMEN Herbst, D. R. 1030 Kauai US 4/7/1968
168 Zanthoxylum dipetalum var. dipetalum mountains in vicinity of kaholuamanu. PRESERVED_SPECIMEN c. n. forbes 347.k Kauai US
169 Zanthoxylum dipetalum var. dipetalum Kalalau Valley, drainage west of Pohakulele Camp and east of Huna. Mixed mesic forest PRESERVED_SPECIMEN Wood, K. R.; DeMotta, M. 11655 Kauai US 12/15/2005
170 Zanthoxylum dipetalum var. dipetalum koolau range, pupukea-paumalu forest reserve, pupukea road, slope below trail PRESERVED_SPECIMEN r. s. cowan 774 Oahu US 1/25/1948
171 Zanthoxylum dipetalum var. dipetalum kipuka puaulu. PRESERVED_SPECIMEN j. f. rock & -- copeland 12572 Hawaii US
172 Zanthoxylum dipetalum var. dipetalum dupont trail, northern slope of kaala PRESERVED_SPECIMEN o. degener, h. clay & a. b. greenwell 19431 Oahu US 2/22/1949
173 Zanthoxylum dipetalum var. dipetalum halemanu. PRESERVED_SPECIMEN d. r. herbst 1030 Kauai US 4/7/1968
174 Zanthoxylum dipetalum var. dipetalum Waimea District, Kawaiula Valley, Kuia Natural Area Reserve. Mesic forest PRESERVED_SPECIMEN Wood, K. R.; Query, M. 14967 Kauai US 5/9/2012
175 Zanthoxylum dipetalum var. dipetalum Lahaina District, W. Maui, Kauaula Valley, south slope, about 1/3 down from valley rim towards stream. PRESERVED_SPECIMEN Oppenheimer, H. L.; Bustamente, K. M.; Perlman, S. P. H21405 Maui US 2/13/2014
176 Zanthoxylum dipetalum var. dipetalum waimea district, nuololo, north facing slopes above northern branch. PRESERVED_SPECIMEN k. r. wood, m. query & m. kirkpatrick 15098 Hawaii US 7/18/2012
177 Zanthoxylum dipetalum var. dipetalum Kipuka Puaulu. PRESERVED_SPECIMEN Rock, J. F.; Copeland, -- 12572 Hawaii US 7/15/2020
178 Zanthoxylum dipetalum var. dipetalum hanalei district; pohakuao; hanging valley between kalalau and hanakoa, below puu ki; nw aspect PRESERVED_SPECIMEN k. r. wood, s. p. perlman & j. lau 1769 Kauai US 4/2/1992
179 Zanthoxylum dipetalum var. dipetalum Waianae Range, Pu'u Ka'ala, above Waialua on the DuPonte trail PRESERVED_SPECIMEN Stone, B. C. 3185 Oahu US 12/20/1959
180 Zanthoxylum dipetalum var. dipetalum kokee. awa'awapuhiu trail. In forest. PRESERVED_SPECIMEN b. c. stone 3338 Kauai US 4/12/1960
181 Zanthoxylum dipetalum var. dipetalum kalalau, between first and second kalalau lookout. PRESERVED_SPECIMEN k. r. wood 11128 Oahu US 2/8/2005
182 Zanthoxylum dipetalum var. dipetalum Puu Kaua, Waianae Mts. PRESERVED_SPECIMEN St. John, H. 20125 Oahu US 12/1/1940
183 Zanthoxylum dipetalum var. dipetalum makai of puu moa PRESERVED_SPECIMEN o. degener, -- salucop & arlantico 11926 Oahu US 4/3/1938
184 Zanthoxylum dipetalum var. dipetalum Hanalei District; Pohakuao; Hanging valley between Kalalau and Hanakoa, below Puu Ki; NW aspect PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P.; Lau, J. 1769 Kauai US 4/2/1992
185 Zanthoxylum dipetalum var. dipetalum Haupu summit region, steep upper southwestern ridges. PRESERVED_SPECIMEN K. R. Wood & M. Wood 11973 Kauai US 7/14/2006
186 Zanthoxylum dipetalum var. dipetalum Lahaina District, Kauaula Valley, south slope, below Helu summit, West Maui. Mesic forest PRESERVED_SPECIMEN S. P. Perlman, H. L. Oppenheimer & K. M. Bustamente 23854 Maui US 2/13/2014
187 Zanthoxylum dipetalum var. dipetalum Kalahu, W of Kalalau Valley PRESERVED_SPECIMEN Degener, O.; Hatheway, W. H. 20596 Kauai US 2/1/1950
188 Zanthoxylum dipetalum var. dipetalum honopu trail Beside trail PRESERVED_SPECIMEN marr, k.l. collector number: km 14 Kauai BISH 3/26/1988
189 Zanthoxylum dipetalum var. dipetalum halemanu, kokee Open woods PRESERVED_SPECIMEN degener, o. degener, i. collector number: 28099 Kauai BISH 12/2/1960
190 Zanthoxylum dipetalum var. dipetalum puu kaua, waianae mts. PRESERVED_SPECIMEN h. st. john 12226 Oahu US 11/6/1932
191 Zanthoxylum dipetalum var. dipetalum Koolau Range, Pupukea-Kahuku trail, insmall ravine near road PRESERVED_SPECIMEN Pearsall, G. A. 522 Oahu US 6/18/1960
192 Zanthoxylum dipetalum var. dipetalum kawaihae uka PRESERVED_SPECIMEN hillebrand, w.b. collector number: s.n. Hawaii BISH
193 Zanthoxylum dipetalum var. dipetalum Kalalau, valley slopes below Puu o Kila. Metrosideros mixed mesic forest adjacent to mesic cliffs PRESERVED_SPECIMEN K. R. Wood 18298 Kauai PTBG 8/28/2019
194 Zanthoxylum dipetalum var. dipetalum lihue-koloa fr, ridge ese of puu kolo In Metrosideros-Acacia koa-Sapindus saponaria mesic forest. PRESERVED_SPECIMEN flynn, t. kawakami, g.; nagata, r. collector number: 3149 Hawaii BISH 9/21/1988
195 Zanthoxylum dipetalum var. dipetalum HAWAII: Puuwaawaa near SE corner; 130 deg. and 300 m SE of cabin; south of road; lowland forest with Myrsine, Sophora chrysophylla, Chamaesyce olowaluana. Threats: cattle, ants, Pennisetm setaceaum, Grevellia robusta. Elev. 3670 aa lava with large surface boulders and rich brown soil PRESERVED_SPECIMEN K. R. Wood 7770 Hawaii PTBG 4/7/1999
196 Zanthoxylum dipetalum var. dipetalum puaulu PRESERVED_SPECIMEN rock, j.f. 10208 BRU
197 Zanthoxylum dipetalum var. dipetalum kipuka puaulu PRESERVED_SPECIMEN rock, j.f.c. collector number: 12572 Hawaii BISH 7/1/1915
198 Zanthoxylum dipetalum var. dipetalum w waimea drainage basin PRESERVED_SPECIMEN forbes, c.n. collector number: 1098 k Kauai BISH 7/3/1917
199 Zanthoxylum dipetalum var. dipetalum hanalei district, upper limahuli preserve, near upper weather port. wainiha side. PRESERVED_SPECIMEN m. edmonds & m. chandler me76 Kauai US 1/25/2012
200 Zanthoxylum dipetalum var. dipetalum Hanalei District, Upper Limahuli Preserve, near upper weather port. Wainiha side. PRESERVED_SPECIMEN M. Edmonds & M. Chandler ME76 Kauai US 1/25/2012
201 Zanthoxylum dipetalum var. dipetalum Koolau Range, Pupukea-Kahuku Trail, Paumalu-Waimea Ridge. PRESERVED_SPECIMEN St. John, H. 11585 Oahu US 3/6/1932
202 Zanthoxylum dipetalum var. dipetalum mohihi rd. kauaikinana stream, ~20 yds. below dam PRESERVED_SPECIMEN flynn, t. collector number: 438 Kauai BISH 6/2/1983
203 Zanthoxylum dipetalum var. dipetalum Waimea District, Kawaiula Valley, Kuia Natural Area Reserve. Mesic forest PRESERVED_SPECIMEN K. R. Wood & M. Query 14967 Kauai US 5/9/2012
204 Zanthoxylum dipetalum var. dipetalum Kalalau, very back on East side of Valley, 50 ft up talus slope on East side of N running gulch, gulch just West of the mainstream. Old relic mixed mesic forest with weedy understory. PRESERVED_SPECIMEN Natalia Tangalin 2385 Kauai PTBG 8/22/2010
205 Zanthoxylum dipetalum var. dipetalum Haupu summit region, steep upper southwestern ridges. PRESERVED_SPECIMEN Wood, K. R.; Wood, M. M. 11973 Kauai US 7/14/2006
206 Zanthoxylum dipetalum var. dipetalum n kaala, dupont trail forest PRESERVED_SPECIMEN degener, o. clay, h.f.; greenwell, a. collector number: 19431 Oahu BISH 2/22/1949
207 Zanthoxylum dipetalum var. dipetalum Dupont Trail, northern slope of Kaala PRESERVED_SPECIMEN Degener, O.; Clay, H. F.; Greenwell, A. B. 19431 Oahu US 2/22/1949
208 Zanthoxylum dipetalum var. dipetalum kumuwela ridge PRESERVED_SPECIMEN degener, o. degener, i. collector number: 28094 Hawaii BISH 9/14/1955
209 Zanthoxylum dipetalum var. dipetalum kawaikoa stream trail, above trail on s side of stream 70 m before .5 mi marker PRESERVED_SPECIMEN marr, k.l. collector number: km 46 Kauai BISH 6/15/1988
210 Zanthoxylum dipetalum var. dipetalum Oio, Paumalu Trail PRESERVED_SPECIMEN Degener, O. 17205 Oahu US 6/16/1940
211 Zanthoxylum dipetalum var. dipetalum koolau range, pupukea-paumalu forest preserve, pupukea road In moist place PRESERVED_SPECIMEN cowan, r.s. collector number: 774 Oahu BISH 1/25/1948
212 Zanthoxylum dipetalum var. dipetalum pupukea military trail PRESERVED_SPECIMEN macdaniels, l.h. collector number: 530 Oahu BISH 1/29/1927
213 Zanthoxylum dipetalum var. dipetalum waianae range. south palawai gulch PRESERVED_SPECIMEN wilbur, r.l. collector number: 620 Oahu BISH 3/28/1948
214 Zanthoxylum dipetalum var. dipetalum ridge .75 mi n of laaukahi, koloa scrubby slope PRESERVED_SPECIMEN st.john, h. cowan, r.s.; webster, g.l.; wilbur, r.l. collector number: 23011 Maui BISH 12/24/1947
215 Zanthoxylum dipetalum var. dipetalum Kaholuamano PRESERVED_SPECIMEN Hitchcock, A. S. 15296 Kauai US 10/20/2016
216 Zanthoxylum dipetalum var. dipetalum kalahu, w of kalalau valley PRESERVED_SPECIMEN o. degener & w. h. hatheway 20596 Kauai US 2/1/1950
217 Zanthoxylum dipetalum var. dipetalum kokee, awaawapuhi trail, ~40 yds from main trail, at #37 PRESERVED_SPECIMEN flynn, t. collector number: 395 Kauai BISH 4/19/1983
218 Zanthoxylum dipetalum var. dipetalum Plateau just S of Awaawapuhi stream head Diverse mesic forest intermixed with open patches of Dicranopteris. With Metrosideros, Ilex, Scaevola sp., Dodonaea, Pouteria, Psychotria greenwelliae, Melicope anisata, Coprosma, Psidium cattleianum, Carex meyenii, Alyxia stellata, and occasional Hedychium. PRESERVED_SPECIMEN Trauernicht, C. Tangalin, N. Collector Number: 750 Kauai BISH 2/19/2009
219 Zanthoxylum dipetalum var. dipetalum puu kaua, waianae mts, steep east ridge PRESERVED_SPECIMEN st.john, h. collector number: 20125 Oahu BISH 12/1/1940
220 Zanthoxylum dipetalum var. dipetalum kipuka puaulu PRESERVED_SPECIMEN rock, j.f.c. hashimoto, t. collector number: 17152 Hawaii BISH 7/1/1918
221 Zanthoxylum dipetalum var. dipetalum kilauea, kipuka puaulu PRESERVED_SPECIMEN rock, j.f.c. collector number: 5916 BISH 12/1/1914
222 Zanthoxylum dipetalum var. dipetalum puu kaua, s ekahanui, subgulch PRESERVED_SPECIMEN perlman, s.p. obata, j. collector number: 6113 Niihau BISH 6/28/1987
223 Zanthoxylum dipetalum var. dipetalum kalahu, w of kalalau valley Open Dicranopteris forest PRESERVED_SPECIMEN degener, o. hatheway, w.h. collector number: 20596 Kauai BISH 2/1/1950
224 Zanthoxylum dipetalum var. dipetalum kokee, halemanu rd, 30 m past turnoff at kokee state park sign PRESERVED_SPECIMEN marr, k.l. collector number: km 42 Kauai BISH 6/13/1988
225 Zanthoxylum dipetalum var. dipetalum kawaikoi stream, by sugi grove Along bank of stream. PRESERVED_SPECIMEN corn, c. collector number: s.n. Kauai BISH 8/29/1982
226 Zanthoxylum dipetalum var. dipetalum Waianae Mts., Honouliuli FR, ridge to Puu Kaua exposed arid cliff PRESERVED_SPECIMEN Herbst, D.R. 5263 Oahu BISH 3/16/1975
227 Zanthoxylum dipetalum var. dipetalum hvnp, kipuka puaulu, next to nps transect Sapling growing beneath KM 19. PRESERVED_SPECIMEN marr, k.l. collector number: km 91 Hawaii BISH 8/1/1988
228 Zanthoxylum dipetalum var. dipetalum Waimea District; Kawaiiki Zone, Below Kalukaula Ridge, on ridge looking down into drainage of Koaie Canyon and north of Waialae. Degraded Metrosideros - Acacia koa Diverse Mesic Forest Myoporum, Cryptocarya, Kadua affinis, Alyxia, Claoxylon sandwicensis, Dodonea, Leptocaphylla, Dryopteris, Zanthoxylum, Dubautia, Psychotria mariniana, Scaevola procera, Melicope PRESERVED_SPECIMEN Tangalin, N. 2573 Kauai BISH 3/23/2011
229 Zanthoxylum dipetalum var. dipetalum w kaunala PRESERVED_SPECIMEN russ, g.w. collector number: 706657 Oahu BISH 5/24/1933
230 Zanthoxylum dipetalum var. dipetalum kamiloloa, ridge above gulch dry; native trees and shrub; exotic grass and shrubs PRESERVED_SPECIMEN stemmermann, r.l. montgomery, s. collector number: 3919 Molokai BISH 7/15/1979
231 Zanthoxylum dipetalum var. dipetalum Anahola Mountains, near SE end of range. Mesic forest on ridge. W/ Metrosideros, Dicranopteris, Carex, Chrysodracon, Psydrax, Machaerina mariscoides. PRESERVED_SPECIMEN Edmonds, M. ME309 Kauai BISH 3/22/2015
232 Zanthoxylum dipetalum var. dipetalum honopu trail Trail-side PRESERVED_SPECIMEN marr, k.l. nakamura, j. collector number: km 40 Kauai BISH 6/11/1988
233 Zanthoxylum dipetalum var. dipetalum kauaikinana stream, just below diversion dam PRESERVED_SPECIMEN marr, k.l. collector number: km 13 Hawaii BISH 3/26/1988
234 Zanthoxylum dipetalum var. dipetalum ridge, puu kaua, honouliuli fr Metrosideros, Acacia koa, grass PRESERVED_SPECIMEN nagata, k.m. collector number: 1156 Oahu BISH 6/11/1973
235 Zanthoxylum dipetalum var. dipetalum kilauea volcano, kipuka puaulu dry stream bed PRESERVED_SPECIMEN rock, j.f.c. collector number: 1020[8] Oahu BISH 7/1/1911
236 Zanthoxylum dipetalum var. dipetalum honopu trail PRESERVED_SPECIMEN marr, k.l. nakamura, j. collector number: km 39 Kauai BISH 6/11/1988
237 Zanthoxylum dipetalum var. dipetalum bird park (kipuka puaulu) moderate forest PRESERVED_SPECIMEN morley, h. collector number: 104-h Hawaii BISH 8/11/1934
238 Zanthoxylum dipetalum var. dipetalum kilauea volcano, kipuka puaulu PRESERVED_SPECIMEN rock, j.f.c. collector number: 1020[8] Hawaii BISH 7/1/1911
239 Zanthoxylum dipetalum var. dipetalum makaha rd, 1.5 m down valley, in sm valley n of kakio keakeo picnic area N facing slope PRESERVED_SPECIMEN marr, k.l. collector number: km 12 Kauai BISH 3/26/1988
240 Zanthoxylum dipetalum var. dipetalum kauai PRESERVED_SPECIMEN knudsen collector number: 82 Molokai BISH
241 Zanthoxylum dipetalum var. dipetalum halemanu Growing in disturbed open part of Acacia-Metrosideros forest with mixed understory. With Pipturus, Coprosma, Physalis peruviana, thimbleberry & ferns. Also grows in pasture of ranch. PRESERVED_SPECIMEN takeuchi, w.n. pyle, l. collector number: 3672 Hawaii BISH 9/26/1987
242 Zanthoxylum dipetalum var. dipetalum n haupu moist PRESERVED_SPECIMEN macdaniels, l.h. collector number: 737 Kauai BISH 2/16/1927
243 Zanthoxylum dipetalum var. dipetalum puu kana PRESERVED_SPECIMEN topping, d.l. collector number: 3286 Oahu BISH 3/21/1926
244 Zanthoxylum dipetalum var. dipetalum Kokee. App. 30m past Kokee State Park sign on right side of main road. Kokee State Park PRESERVED_SPECIMEN Marr, K.L. K.M. 15 Kauai BISH 3/26/1988
245 Zanthoxylum dipetalum var. dipetalum halemanu-kokee trail, .5 mi fm upper parking lot. upper end of gully, s sside of trail 20 m fm trail where it levels off dry, open PRESERVED_SPECIMEN marr, k.l. nakamura, j. collector number: km 43 Oahu BISH 6/13/1988
246 Zanthoxylum dipetalum var. dipetalum e of kumuweia ridge, kokee Open forest along ditch trail PRESERVED_SPECIMEN degener, o. collector number: 20598 Kauai BISH 2/7/1950
247 Zanthoxylum dipetalum var. dipetalum halemanu Dry-mesic forest. PRESERVED_SPECIMEN herbst, d.r. collector number: 1030 Kauai BISH 4/7/1968
248 Zanthoxylum dipetalum var. dipetalum puu kaua, waianae mts. honouliuli wooded steep east ridge PRESERVED_SPECIMEN st.john, h. collector number: 12226 Oahu BISH 11/6/1932
249 Zanthoxylum dipetalum var. dipetalum ridge, kaua to kanehoa Psychotria hathewayi PRESERVED_SPECIMEN takeuchi, w.n. Oahu BISH 10/1/1986
250 Zanthoxylum dipetalum var. dipetalum kaholuamano Myoporum and lava field PRESERVED_SPECIMEN rock, j.f.c. collector number: 2115 Hawaii BISH 3/3/1909
251 Zanthoxylum dipetalum var. dipetalum kawaikoa stream trail, above trail on s side of stream 70 m before .5 mi marker PRESERVED_SPECIMEN marr, k.l. collector number: km 45 Kauai BISH 6/15/1988
252 Zanthoxylum dipetalum var. dipetalum waimea canyon rim, below road, 11.8 mi fm waimea PRESERVED_SPECIMEN marr, k.l. collector number: km 10 Kauai BISH 3/26/1988
253 Zanthoxylum dipetalum var. dipetalum kumuweia ridge, e of waineke swamp PRESERVED_SPECIMEN degener, o. greenwell, a. collector number: 21551 Oahu BISH 1/8/1952
254 Zanthoxylum dipetalum var. dipetalum kipuka puaulu PRESERVED_SPECIMEN rock, j.f.c. Hawaii BISH 8/8/1917
255 Zanthoxylum dipetalum var. dipetalum puu kaua, ridge, about 2200' PRESERVED_SPECIMEN suehiro, a. Oahu BISH 11/6/1932
256 Zanthoxylum dipetalum var. dipetalum kokee, milolii rd, 0.35 mi. after jct. with makaha rd PRESERVED_SPECIMEN marr, k.l. collector number: 18 Maui BISH 3/26/1988
257 Zanthoxylum dipetalum var. dipetalum upper limahuli valley PRESERVED_SPECIMEN tangalin, n. collector number: 898 BISH 8/8/2006
258 Zanthoxylum dipetalum var. dipetalum waimea district; kawaiula valley, kuia natural area reserve Metrosideros-Acacia mesic forest with Alphitonia ponderosa, Diospyros sandwicensis, dracaena aurea, Psychotria mariniana, Dodonaea viscosa, Wikstroemia furcata, Planchnella sandwicensis, dryopteris glabra, Doodia kunthiana, Panicum PRESERVED_SPECIMEN wood, k.r. query, m. collector number: 14967 Kauai BISH 5/9/2012
259 Zanthoxylum dipetalum var. dipetalum waimea, 1.3 mi. down makaha ridge road from hwy. 550, below kokio kookeo picnic area Exposed at low tide on limestone benches east of Cove. PRESERVED_SPECIMEN flynn, t. uchida, g. collector number: 2773 Oahu BISH 2/24/1988
260 Zanthoxylum dipetalum var. dipetalum waianae mts., s ekanui gulch, ridge in shaded site under Eucalyptus PRESERVED_SPECIMEN obata, j.k. palmer, d.; funk, e. collector number: 394 Maui BISH 7/9/1978
261 Zanthoxylum dipetalum var. dipetalum 2 ridges s. of pohakea pass forest PRESERVED_SPECIMEN christophersen, e. wilder, g.p.; hume, e.p. collector number: 1658 Oahu BISH 3/19/1931
262 Zanthoxylum dipetalum var. dipetalum kalalau valley, s rim and upper reaches, 0.4 miles e of kalalau lookout; steep slopes with low mixed mesophytic forest of Metrosideros, Pelea, Xylosma & Pleomele PRESERVED_SPECIMEN lorence, d.h. flynn, t.; de lappe, r.; chong, d. collector number: 5121 Kauai BISH 3/20/1987
263 Zanthoxylum dipetalum var. dipetalum halemanu PRESERVED_SPECIMEN takeuchi, w.n. pyle, l. collector number: 3678 Lanai BISH 9/26/1987
264 Zanthoxylum dipetalum var. dipetalum awaawapuhi trail; honopu Moist Acacia Forest. PRESERVED_SPECIMEN lane, i.e. collector number: 50-625 Kauai BISH 12/29/1956
265 Zanthoxylum dipetalum var. dipetalum puu kaua, waianae mts., steep e. ridge PRESERVED_SPECIMEN st.john, h. collector number: 20125 Oahu BISH 12/1/1940
266 Zanthoxylum dipetalum var. dipetalum makakupaia, s fork, kaunakakai gulch Xylosma, Coprosma, Hedychium, Osmanthus, Wikstroemia, Melinis, Microlepia PRESERVED_SPECIMEN lau, j. collector number: 1465 Molokai BISH 9/25/1985
267 Zanthoxylum dipetalum var. dipetalum kumuwela ridge, waimea Dry open woods PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13822 Kauai BISH 12/28/1933
268 Zanthoxylum dipetalum var. dipetalum pupukea paumalu fr, off jeep road PRESERVED_SPECIMEN landgraf, m.f. Oahu BISH 5/18/1967
269 Zanthoxylum dipetalum var. dipetalum plateau just s of awaawapuhi stream head Diverse mesic forest intermixed with open patches of Dicranopteris. With Metrosideros, Ilex, Scaevola sp., Dodonaea, Pouteria, Psychotria greenwelliae, Melicope anisata, Coprosma, Psidium cattleianum, Carex meyenii, Alyxia stellata, and occasional Hedychium. PRESERVED_SPECIMEN trauernicht, c. tangalin, n. collector number: 750 Kauai BISH 2/19/2009
270 Zanthoxylum dipetalum var. dipetalum hikimoe, n facing slopes below road Small patches of Diospyros sandwicensis forest with occasional Metrosideros polymorpha, Coprosma waimeae, Psychotria greenwelliae, Zanthoxylum dipetelum, Psydrax odorata, Pouteria sandwicensis, Nestegis sandwicensis, Pleomele aurea, Pisonia sp., Syzygium sandwicensis, gulch is mostly dominated by secondary invasive species. PRESERVED_SPECIMEN wood, k.r. perlman, s. collector number: 14226 Kauai BISH 6/1/2010
271 Zanthoxylum dipetalum var. dipetalum koloa, haupu open shrubland, herbland, sparse tree cover; Metrosideros polymorpha, Dicranopteris linearis, Cibotium glaucum, Sadleria pallida, Hedyotis terminalis, Psychotria mariniana, Antidesma platyphyllum, Bobea brevipes, Xylosma hawaiiense, Cheirodendron fauriei, Melicope feddei, Tetraplasandra PRESERVED_SPECIMEN wood, k.r. wood, m. collector number: 11973 Kauai BISH 7/14/2006
272 Zanthoxylum dipetalum var. dipetalum hvnp, kipuka puaulu. next to nps transect PRESERVED_SPECIMEN drake, d. collector number: k.m. 19 Maui BISH 3/25/1988
273 Zanthoxylum dipetalum var. dipetalum hanalei, hoolulu valley Nr ridge crest in mixed mesic forest PRESERVED_SPECIMEN lau, j. collector number: 2205 Kauai BISH 4/1/1986
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